European islands, development and the cohesion policy : a case study of Kokar, Aland islands

A European Union initiative is seeking ways of determining the development potential of Europe’s lagging regions, which include various islands. On the basis of the policy review, methodology and data collected by the ongoing EUROISLANDS project, this paper presents Kökar (population: 262), the easternmost municipality of the Åland Islands, as a prototype archipelago that is seriously challenged by its size and multiple peripherality. It reviews the state of its social and transportation infrastructure, and of its human and financial capital. While the situation is serious, there are opportunities for branding, for developing associated economic activity, as well for appealing to a new wave of residents.peer-reviewe

Nephtyidae (Annelida, Polychaeta) from southern Europe

Eighteen nephtyid species belonging to four different genera are known from southern Europe. In this study we revise the nephtyids from this area and provide descriptions of all the known species. Geographical and bathymetrical distributions are given, together with ecological notes. Inermonephtys foretmontardoi, new species, is described, Micronephthys maryae is newly synonymized with M. stammeri and Aglaophamus rubellus with A. agilis. New diagnoses are provided for southern European genera and a key to all species from the region is included.FCT - SFRH/BD/16563/2004EC FP6 - GOCE-CT-2005-511234 HERMESEC FP7 - PROJ 226354 HERMIONEVR, dnr 2009-512

Have ozone effects on carbon sequestration been overestimated?: a new biomass response function for wheat

Elevated levels of tropospheric ozone can significantly impair the growth of crops. The reduced removal of CO2 by plants leads to higher atmospheric concentrations of CO2, enhancing radiative forcing. Ozone effects on economic yield, e.g. the grain yield of wheat (Triticum aestivum L.), are currently used to model effects on radiative forcing. However, changes in grain yield do not necessarily reflect changes in total biomass. Based on an analysis of 22 ozone exposure experiments with field-grown wheat, we investigated whether the use of effects on grain yield as a proxy for effects on biomass under- or overestimates effects on biomass. First, we confirmed that effects on partitioning and biomass loss are both of significant importance for wheat yield loss. Then we derived ozone dose response functions for biomass loss and for harvest index (the proportion of above-ground biomass converted to grain) based on 12 experiments and recently developed ozone uptake modelling for wheat. Finally, we used a European-scale chemical transport model (EMEP MSC-West) to assess the effect of ozone on biomass (−9%) and grain yield (−14%) loss over Europe. Based on yield data per grid square, we estimated above-ground biomass losses due to ozone in 2000 in Europe, totalling 22.2 million tonnes. Incorrectly applying the grain yield response function to model effects on biomass instead of the biomass response function of this paper would have indicated total above-ground biomass losses totalling 38.1 million (i.e. overestimating effects by 15.9 million tonnes). A key conclusion from our study is that future assessments of ozone-induced loss of agroecosystem carbon storage should use response functions for biomass, such as that provided in this paper, not grain yield, to avoid overestimation of the indirect radiative forcing from ozone effects on crop biomass accumulation

The Relationship Between Birch Pollen, Air Pollution and Weather Types and Their Effect on Antihistamine Purchase in Two Swedish Cities

Exposure to elevated air pollution levels can aggravate pollen allergy symptoms. The aim of this study was to investigate the relationships between airborne birch (Betula) pollen, urban air pollutants NO2, O3 and PM10 and their effects on antihistamine demand in Gothenburg and Malmo¨, Sweden, 2006–2012. Further, the influence of large-scale weather pattern on pollen-/ pollution-related risk, using Lambweather types (LWTs), was analysed. Daily LWTs were obtained by comparing the atmospheric pressure over a 16-point grid system over southern Sweden (scale *3000 km). They include two non-directional types, cyclonic (C) and anticyclonic (A) and eight directional types depending on the wind direction (N, NE, E…). Birch pollen levels were exceptionally high under LWTs E and SE in both cities. Furthermore, LWTs with dry and moderately calm meteorological character (A, NE, E, SE) were associated with strongly elevated air pollution (NO2 and PM10) in Gothenburg. For most weather situations in both cities, simultaneously high birch pollen together with high air pollution had larger over-the-counter (OTC) sales of antihistamines than situations with high birch pollen alone. LWTs NE, E, SE and S had the highest OTC sales in both cities. InGothenburg, the city with a higher load of both birch pollen and air pollution, the higher OTC sales were especially obvious and indicate an increased effect on allergic symptoms from air pollution. Furthermore, GothenburgLWTs A, NE, E and SEwere associatedwith high pollen and air pollution levels and thus classified as high-risk weather types. In Malmo¨, corresponding highrisk LWTs were NE, E, SE and S. Furthermore, occurrence of high pollen and air pollutants as well as OTC sales correlated strongly with vapour pressure deficit and temperature in Gothenburg (much less so in Malmo¨). This provides evidence that the combination of meteorological properties associated with LWTs can explain high levels of birch pollen and air pollution. Our study shows that LWTs represent a useful tool for integrated daily air quality forecasting/warning

Accumulation of antimony and lead in leaves and needles of trees: The role of traffic emissions

Antimony (Sb) is a toxic metalloid, which has been increasingly used in the brake lining of ve-hicles, and increased concentrations have been found in soils near abundant traffic. However, since very few investigations of Sb accumulation in urban vegetation have been undertaken there exists a knowledge gap. We studied the concentrations of Sb in leaves and needles of trees in the Gothenburg City area, Sweden. In addition, lead (Pb), also associated with traffic, was investi-gated. Sb and Pb concentrations of Quercus palustris leaves at seven sites with contrasting traffic intensity varied substantially, correlated with the traffic-related PAH (polycyclic aromatic hy-drocarbon) air pollution at the sites and increased during the growing season. Sb but not Pb concentrations were significantly higher in needles of Picea abies and Pinus sylvestris near major roads compared to sites at larger distances. In Pinus nigra needles at two urban streets both Sb and Pb were higher compared to an urban nature park environment, emphasising the role of traffic emissions for these elements. A continued accumulation of Sb and Pb in three years old needles of Pinus nigra, two years old needles of Pinus sylvestris and eleven years old needles of Picea abies was observed. Our data suggest a pronounced link between traffic pollution and Sb accumulation in leaves and needles, where the particles carrying Sb seem not to be transported very far from the source. We also conclude that there exists a strong potential for Sb and Pb bioaccumulation over time in leaves and needles. Implications of these findings are that increased concentrations of toxic Sb and Pb are likely to prevail in environments with high traffic intensity and that Sb can enter the ecological food chain by accumulation in leaves and needles, which is important for the biogeochemical cycling

Spaghetti to a Tree:A Robust Phylogeny for Terebelliformia (Annelida) Based on Transcriptomes, Molecular and Morphological Data

Terebelliformia—“spaghetti worms” and their allies—are speciose and ubiquitous marine annelids but our understanding of how their morphological and ecological diversity evolved is hampered by an uncertain delineation of lineages and their phylogenetic relationships. Here, we analyzed transcriptomes of 20 terebelliforms and an outgroup to build a robust phylogeny of the main lineages grounded on 12,674 orthologous genes. We then supplemented this backbone phylogeny with a denser sampling of 121 species using five genes and 90 morphological characters to elucidate fine-scale relationships. The monophyly of six major taxa was supported: Pectinariidae, Ampharetinae, Alvinellidae, Trichobranchidae, Terebellidae and Melinninae. The latter, traditionally a subfamily of Ampharetidae, was unexpectedly the sister to Terebellidae, and hence becomes Melinnidae, and Ampharetinae becomes Ampharetidae. We found no support for the recently proposed separation of Telothelepodidae, Polycirridae and Thelepodidae from Terebellidae. Telothelepodidae was nested within Thelepodinae and is accordingly made its junior synonym. Terebellidae contained the subfamily-ranked taxa Terebellinae and Thelepodinae. The placement of the simplified Polycirridae within Terebellinae differed from previous hypotheses, warranting the division of Terebellinae into Lanicini, Procleini, Terebellini and Polycirrini. Ampharetidae (excluding Melinnidae) were well-supported as the sister group to Alvinellidae and we recognize three clades: Ampharetinae, Amaginae and Amphicteinae. Our analysis found several paraphyletic genera and undescribed species. Morphological transformations on the phylogeny supported the hypothesis of an ancestor that possessed both branchiae and chaetae, which is at odds with proposals of a “naked” ancestor. Our study demonstrates how a robust backbone phylogeny can be combined with dense taxon coverage and morphological traits to give insights into the evolutionary history and transformation of traits

Avoided intersections of nodal lines

We consider real eigen-functions of the Schr\"odinger operator in 2-d. The nodal lines of separable systems form a regular grid, and the number of nodal crossings equals the number of nodal domains. In contrast, for wave functions of non integrable systems nodal intersections are rare, and for random waves, the expected number of intersections in any finite area vanishes. However, nodal lines display characteristic avoided crossings which we study in the present work. We define a measure for the avoidance range and compute its distribution for the random waves ensemble. We show that the avoidance range distribution of wave functions of chaotic systems follow the expected random wave distributions, whereas for wave functions of classically integrable but quantum non-separable wave functions, the distribution is quite different. Thus, the study of the avoidance distribution provides more support to the conjecture that nodal structures of chaotic systems are reproduced by the predictions of the random waves ensemble.Comment: 12 pages, 4 figure

A lower bound for nodal count on discrete and metric graphs

According to a well-know theorem by Sturm, a vibrating string is divided into exactly N nodal intervals by zeros of its N-th eigenfunction. Courant showed that one half of Sturm's theorem for the strings applies to the theory of membranes: N-th eigenfunction cannot have more than N domains. He also gave an example of a eigenfunction high in the spectrum with a minimal number of nodal domains, thus excluding the existence of a non-trivial lower bound. An analogue of Sturm's result for discretizations of the interval was discussed by Gantmacher and Krein. The discretization of an interval is a graph of a simple form, a chain-graph. But what can be said about more complicated graphs? It has been known since the early 90s that the nodal count for a generic eigenfunction of the Schrodinger operator on quantum trees (where each edge is identified with an interval of the real line and some matching conditions are enforced on the vertices) is exact too: zeros of the N-th eigenfunction divide the tree into exactly N subtrees. We discuss two extensions of this result in two directions. One deals with the same continuous Schrodinger operator but on general graphs (i.e. non-trees) and another deals with discrete Schrodinger operator on combinatorial graphs (both trees and non-trees). The result that we derive applies to both types of graphs: the number of nodal domains of the N-th eigenfunction is bounded below by N-L, where L is the number of links that distinguish the graph from a tree (defined as the dimension of the cycle space or the rank of the fundamental group of the graph). We also show that if it the genericity condition is dropped, the nodal count can fall arbitrarily far below the number of the corresponding eigenfunction.Comment: 15 pages, 4 figures; Minor corrections: added 2 important reference

Nodal domains statistics - a criterion for quantum chaos

We consider the distribution of the (properly normalized) numbers of nodal domains of wave functions in 2-$d$ quantum billiards. We show that these distributions distinguish clearly between systems with integrable (separable) or chaotic underlying classical dynamics, and for each case the limiting distribution is universal (system independent). Thus, a new criterion for quantum chaos is provided by the statistics of the wave functions, which complements the well established criterion based on spectral statistics.Comment: 4 pages, 5 figures, revte